BLASTP 2.2.12 [Aug-07-2005] Reference: Altschul, Stephen F., Thomas L. Madden, Alejandro A. Schaffer, Jinghui Zhang, Zheng Zhang, Webb Miller, and David J. Lipman (1997), "Gapped BLAST and PSI-BLAST: a new generation of protein database search programs", Nucleic Acids Res. 25:3389-3402. Query= BGIBMGA000167-TA|BGIBMGA000167-PA|IPR004361|Glyoxalase I, IPR004360|Glyoxalase/bleomycin resistance protein/dioxygenase (173 letters) Database: arabidopsis 28,952 sequences; 12,070,560 total letters Searching..................................................done Score E Sequences producing significant alignments: (bits) Value At1g08110.2 68414.m00889 lactoylglutathione lyase, putative / gl... 221 3e-58 At1g08110.1 68414.m00888 lactoylglutathione lyase, putative / gl... 221 3e-58 At1g67280.1 68414.m07657 lactoylglutathione lyase, putative / gl... 68 4e-12 At1g11840.3 68414.m01360 lactoylglutathione lyase, putative / gl... 66 9e-12 At1g11840.2 68414.m01362 lactoylglutathione lyase, putative / gl... 66 9e-12 At1g11840.1 68414.m01361 lactoylglutathione lyase, putative / gl... 66 9e-12 At2g28420.1 68415.m03453 lactoylglutathione lyase family protein... 33 0.14 At2g32090.2 68415.m03923 lactoylglutathione lyase family protein... 31 0.56 At2g32090.1 68415.m03922 lactoylglutathione lyase family protein... 31 0.56 At4g36030.1 68417.m05129 armadillo/beta-catenin repeat family pr... 30 0.98 At1g15380.1 68414.m01841 lactoylglutathione lyase family protein... 29 2.3 At3g01810.1 68416.m00123 expressed protein 28 3.0 At3g27300.1 68416.m03412 glucose-6-phosphate 1-dehydrogenase / G... 28 3.9 At1g07645.1 68414.m00822 lactoylglutathione lyase family protein... 28 3.9 At5g40760.1 68418.m04946 glucose-6-phosphate 1-dehydrogenase / G... 27 5.2 At5g03150.1 68418.m00263 zinc finger (C2H2 type) family protein ... 27 5.2 At4g21150.1 68417.m03057 ribophorin II (RPN2) family protein con... 27 5.2 At3g22330.1 68416.m02820 DEAD box RNA helicase, putative similar... 27 5.2 At3g06640.1 68416.m00772 protein kinase family protein contains ... 27 9.1 >At1g08110.2 68414.m00889 lactoylglutathione lyase, putative / glyoxalase I, putative similar to lactoylglutathione lyase SP:O04885 from [Brassica juncea] Length = 185 Score = 221 bits (539), Expect = 3e-58 Identities = 99/165 (60%), Positives = 121/165 (73%), Gaps = 2/165 (1%) Query: 6 SPQEIEALCQTPDPTTKDFMFQQTMYRIKDPRKTIPFYTGVLGMTLLKQLHFPAMKFSLF 65 SP L D TK ++ QQTM+RIKDP+ ++ FY+ VLGM+LLK+L F MKFSL+ Sbjct: 8 SPANNPGLSTNRDEATKGYIMQQTMFRIKDPKASLDFYSRVLGMSLLKRLDFSEMKFSLY 67 Query: 66 FMGYENPAEVPRDDDARTKWAMTRKATLELTYNWGTESDD--SSYHNGNSDPRGFGHIGI 123 F+GYE+ P D RT W + AT+ELT+NWGTESD YHNGNS+PRGFGHIG+ Sbjct: 68 FLGYEDTTTAPTDPTERTVWTFGQPATIELTHNWGTESDPEFKGYHNGNSEPRGFGHIGV 127 Query: 124 LVPDVDVACARFEQLGVKFIKRPNDGKMKGLAFIQDPDGYWIEIF 168 V DV AC RFE+LGV+F K+PNDGKMK +AFI+DPDGYWIEIF Sbjct: 128 TVDDVHKACERFEELGVEFAKKPNDGKMKNIAFIKDPDGYWIEIF 172 >At1g08110.1 68414.m00888 lactoylglutathione lyase, putative / glyoxalase I, putative similar to lactoylglutathione lyase SP:O04885 from [Brassica juncea] Length = 185 Score = 221 bits (539), Expect = 3e-58 Identities = 99/165 (60%), Positives = 121/165 (73%), Gaps = 2/165 (1%) Query: 6 SPQEIEALCQTPDPTTKDFMFQQTMYRIKDPRKTIPFYTGVLGMTLLKQLHFPAMKFSLF 65 SP L D TK ++ QQTM+RIKDP+ ++ FY+ VLGM+LLK+L F MKFSL+ Sbjct: 8 SPANNPGLSTNRDEATKGYIMQQTMFRIKDPKASLDFYSRVLGMSLLKRLDFSEMKFSLY 67 Query: 66 FMGYENPAEVPRDDDARTKWAMTRKATLELTYNWGTESDD--SSYHNGNSDPRGFGHIGI 123 F+GYE+ P D RT W + AT+ELT+NWGTESD YHNGNS+PRGFGHIG+ Sbjct: 68 FLGYEDTTTAPTDPTERTVWTFGQPATIELTHNWGTESDPEFKGYHNGNSEPRGFGHIGV 127 Query: 124 LVPDVDVACARFEQLGVKFIKRPNDGKMKGLAFIQDPDGYWIEIF 168 V DV AC RFE+LGV+F K+PNDGKMK +AFI+DPDGYWIEIF Sbjct: 128 TVDDVHKACERFEELGVEFAKKPNDGKMKNIAFIKDPDGYWIEIF 172 >At1g67280.1 68414.m07657 lactoylglutathione lyase, putative / glyoxalase I, putative similar to putative lactoylglutathione lyase SP:Q39366, GI:2494843 from [Brassica oleracea] Length = 350 Score = 67.7 bits (158), Expect = 4e-12 Identities = 51/142 (35%), Positives = 68/142 (47%), Gaps = 29/142 (20%) Query: 30 MYRIKDPRKTIPFYTGVLGMTLLKQLHFPAMKFSLFFMGYENPAEVPRDDDARTKWAMTR 89 +YR+ D +TI FYT LGM LL++ P K++ F+GY P D Sbjct: 93 VYRVGDMDRTIKFYTECLGMKLLRKRDIPEEKYTNAFLGYG-----PEDSHF-------- 139 Query: 90 KATLELTYNWGTESDDSSYHNGNSDPRGFGHIGILVPDVDVACARFEQLGVKFIKRPNDG 149 +ELTYN+G + D GFGH GI V DV + G K + P G Sbjct: 140 --VIELTYNYGVDKYDIG--------AGFGHFGIAVDDVAKTVELVKAKGGKVSREP--G 187 Query: 150 KMKG----LAFIQDPDGYWIEI 167 +KG +AFI+DPDGY E+ Sbjct: 188 PVKGGKTVIAFIEDPDGYKFEL 209 Score = 47.6 bits (108), Expect = 5e-06 Identities = 43/147 (29%), Positives = 59/147 (40%), Gaps = 26/147 (17%) Query: 28 QTMYRIKDPRKTIPFYTGVLGMTLLKQLHFPAMKFSLFFMGYENPAEVPRDDDARTKWAM 87 Q M R+ D + I FY GM LL+ P K+++ MGY P D Sbjct: 221 QVMLRVGDLDRAIKFYEKAFGMELLRTRDNPEYKYTIAMMGYG-----PED--------- 266 Query: 88 TRKATLELTYNWGTESDDSSYHNGNSDPRGFGHIGILVPDVDVACARFEQLGVKFIKRPN 147 + LELTYN+G + Y GN+ + I I DV + G K + P Sbjct: 267 -KFPVLELTYNYGV----TEYDKGNA----YAQIAIGTDDVYKTAEAIKLFGGKITREPG 317 Query: 148 --DGKMKGLAFIQDPDGYWIEIFTSNV 172 G + DPDG W +F N+ Sbjct: 318 PLPGISTKITACLDPDG-WKSVFVDNI 343 >At1g11840.3 68414.m01360 lactoylglutathione lyase, putative / glyoxalase I, putative highly similar to putative lactoylglutathione lyase SP:Q39366 from [Brassica oleracea] Length = 232 Score = 66.5 bits (155), Expect = 9e-12 Identities = 51/151 (33%), Positives = 67/151 (44%), Gaps = 25/151 (16%) Query: 19 PTTKDFMFQQTMYRIKDPRKTIPFYTGVLGMTLLKQLHFPAMKFSLFFMGYENPAEVPRD 78 P + F +YR+ D +TI FYT V GM LL++ P K+S F+G+ P Sbjct: 11 PKKDNRRFLHVVYRVGDLDRTIEFYTEVFGMKLLRKRDIPEEKYSNAFLGF-GPE----- 64 Query: 79 DDARTKWAMTRKATLELTYNWGTESDDSSYHNGNSDPRGFGHIGILVPDVDVACARFEQL 138 T +ELTYN+G SSY G GFGH I DV Sbjct: 65 ---------TSNFVVELTYNYGV----SSYDIGT----GFGHFAISTQDVSKLVENVRAK 107 Query: 139 GVKFIKRPNDGKMKG--LAFIQDPDGYWIEI 167 G + P K G +AF++DPDGY E+ Sbjct: 108 GGNVTREPGPVKGGGSVIAFVKDPDGYTFEL 138 Score = 40.3 bits (90), Expect = 7e-04 Identities = 19/49 (38%), Positives = 27/49 (55%) Query: 26 FQQTMYRIKDPRKTIPFYTGVLGMTLLKQLHFPAMKFSLFFMGYENPAE 74 F Q M R+ D + I FY LGM LL+++ P K+++ MGY E Sbjct: 148 FCQVMLRVGDLDRAIKFYEKALGMRLLRKIERPEYKYTIGMMGYAEEYE 196 >At1g11840.2 68414.m01362 lactoylglutathione lyase, putative / glyoxalase I, putative highly similar to putative lactoylglutathione lyase SP:Q39366 from [Brassica oleracea] Length = 283 Score = 66.5 bits (155), Expect = 9e-12 Identities = 51/151 (33%), Positives = 67/151 (44%), Gaps = 25/151 (16%) Query: 19 PTTKDFMFQQTMYRIKDPRKTIPFYTGVLGMTLLKQLHFPAMKFSLFFMGYENPAEVPRD 78 P + F +YR+ D +TI FYT V GM LL++ P K+S F+G+ P Sbjct: 11 PKKDNRRFLHVVYRVGDLDRTIEFYTEVFGMKLLRKRDIPEEKYSNAFLGF-GPE----- 64 Query: 79 DDARTKWAMTRKATLELTYNWGTESDDSSYHNGNSDPRGFGHIGILVPDVDVACARFEQL 138 T +ELTYN+G SSY G GFGH I DV Sbjct: 65 ---------TSNFVVELTYNYGV----SSYDIGT----GFGHFAISTQDVSKLVENVRAK 107 Query: 139 GVKFIKRPNDGKMKG--LAFIQDPDGYWIEI 167 G + P K G +AF++DPDGY E+ Sbjct: 108 GGNVTREPGPVKGGGSVIAFVKDPDGYTFEL 138 Score = 43.6 bits (98), Expect = 7e-05 Identities = 45/152 (29%), Positives = 63/152 (41%), Gaps = 30/152 (19%) Query: 26 FQQTMYRIKDPRKTIPFYTGVLGMTLLKQLHFPAMKFSLFFMGYENPAEVPRDDDARTKW 85 F Q M R+ D + I FY LGM LL+++ P K+++ MGY E Sbjct: 148 FCQVMLRVGDLDRAIKFYEKALGMRLLRKIERPEYKYTIGMMGYAEEYE----------- 196 Query: 86 AMTRKATLELTYNWGTESDDSSYHNGNSDPRGFGHIGILVPDV----DVACARFEQLGVK 141 LELTYN+ D + Y GN+ + I I DV +V ++LG K Sbjct: 197 ----SIVLELTYNY----DVTEYTKGNA----YAQIAIGTDDVYKSGEVIKIVNQELGGK 244 Query: 142 FIKR--PNDGKMKGLAFIQDPDGYWIEIFTSN 171 + P G + DPDG W + N Sbjct: 245 ITREAGPLPGLGTKIVSFLDPDG-WKTVLVDN 275 >At1g11840.1 68414.m01361 lactoylglutathione lyase, putative / glyoxalase I, putative highly similar to putative lactoylglutathione lyase SP:Q39366 from [Brassica oleracea] Length = 283 Score = 66.5 bits (155), Expect = 9e-12 Identities = 51/151 (33%), Positives = 67/151 (44%), Gaps = 25/151 (16%) Query: 19 PTTKDFMFQQTMYRIKDPRKTIPFYTGVLGMTLLKQLHFPAMKFSLFFMGYENPAEVPRD 78 P + F +YR+ D +TI FYT V GM LL++ P K+S F+G+ P Sbjct: 11 PKKDNRRFLHVVYRVGDLDRTIEFYTEVFGMKLLRKRDIPEEKYSNAFLGF-GPE----- 64 Query: 79 DDARTKWAMTRKATLELTYNWGTESDDSSYHNGNSDPRGFGHIGILVPDVDVACARFEQL 138 T +ELTYN+G SSY G GFGH I DV Sbjct: 65 ---------TSNFVVELTYNYGV----SSYDIGT----GFGHFAISTQDVSKLVENVRAK 107 Query: 139 GVKFIKRPNDGKMKG--LAFIQDPDGYWIEI 167 G + P K G +AF++DPDGY E+ Sbjct: 108 GGNVTREPGPVKGGGSVIAFVKDPDGYTFEL 138 Score = 43.6 bits (98), Expect = 7e-05 Identities = 45/152 (29%), Positives = 63/152 (41%), Gaps = 30/152 (19%) Query: 26 FQQTMYRIKDPRKTIPFYTGVLGMTLLKQLHFPAMKFSLFFMGYENPAEVPRDDDARTKW 85 F Q M R+ D + I FY LGM LL+++ P K+++ MGY E Sbjct: 148 FCQVMLRVGDLDRAIKFYEKALGMRLLRKIERPEYKYTIGMMGYAEEYE----------- 196 Query: 86 AMTRKATLELTYNWGTESDDSSYHNGNSDPRGFGHIGILVPDV----DVACARFEQLGVK 141 LELTYN+ D + Y GN+ + I I DV +V ++LG K Sbjct: 197 ----SIVLELTYNY----DVTEYTKGNA----YAQIAIGTDDVYKSGEVIKIVNQELGGK 244 Query: 142 FIKR--PNDGKMKGLAFIQDPDGYWIEIFTSN 171 + P G + DPDG W + N Sbjct: 245 ITREAGPLPGLGTKIVSFLDPDG-WKTVLVDN 275 >At2g28420.1 68415.m03453 lactoylglutathione lyase family protein / glyoxalase I family protein contains glyoxalase family protein domain, Pfam:PF00903 Length = 184 Score = 32.7 bits (71), Expect = 0.14 Identities = 37/137 (27%), Positives = 53/137 (38%), Gaps = 27/137 (19%) Query: 34 KDPRKTIPFYTGVLGMTLLKQLHFPAMKFSLFFMGYENPAEVPRDDDARTKWAMTRKATL 93 KD +K++ FYT VLG F+ E PA D W + Sbjct: 29 KDVKKSLEFYTKVLG-----------------FVEIERPASFDFDG----AWLFNYGVGI 67 Query: 94 ELTYNWGTESDDSSYHNGNSDPRGFGHIGILVPDVDVACARFEQLGVKFIKRP-NDGKMK 152 L + D + DP HI D++ R +++ VK+IKR D K Sbjct: 68 HLVQ--AKDQDKLPSDTDHLDPMD-NHISFQCEDMEALEKRLKEVKVKYIKRTVGDEKDA 124 Query: 153 GL--AFIQDPDGYWIEI 167 + F DPDG+ +EI Sbjct: 125 AIDQLFFNDPDGFMVEI 141 >At2g32090.2 68415.m03923 lactoylglutathione lyase family protein / glyoxalase I family protein contains glyoxalase family protein domain, Pfam:PF00903 Length = 113 Score = 30.7 bits (66), Expect = 0.56 Identities = 23/69 (33%), Positives = 35/69 (50%), Gaps = 3/69 (4%) Query: 100 GTESDDSSYHNGNSDPRGFGHIGILVPDVDVACARFEQLGVK-FIKRPNDGKMKGLAFIQ 158 G S S+ + + P G HI VP+ D ++ G++ F K DGK+K + F Sbjct: 43 GPYSATSAVKDPSHLPMGH-HICFSVPNFDSFLHSLKEKGIETFQKSLPDGKVKQVFFF- 100 Query: 159 DPDGYWIEI 167 DPDG +E+ Sbjct: 101 DPDGNGLEV 109 >At2g32090.1 68415.m03922 lactoylglutathione lyase family protein / glyoxalase I family protein contains glyoxalase family protein domain, Pfam:PF00903 Length = 135 Score = 30.7 bits (66), Expect = 0.56 Identities = 23/69 (33%), Positives = 35/69 (50%), Gaps = 3/69 (4%) Query: 100 GTESDDSSYHNGNSDPRGFGHIGILVPDVDVACARFEQLGVK-FIKRPNDGKMKGLAFIQ 158 G S S+ + + P G HI VP+ D ++ G++ F K DGK+K + F Sbjct: 65 GPYSATSAVKDPSHLPMGH-HICFSVPNFDSFLHSLKEKGIETFQKSLPDGKVKQVFFF- 122 Query: 159 DPDGYWIEI 167 DPDG +E+ Sbjct: 123 DPDGNGLEV 131 >At4g36030.1 68417.m05129 armadillo/beta-catenin repeat family protein contains Pfam profile: PF00514 armadillo/beta-catenin-like repeat Length = 670 Score = 29.9 bits (64), Expect = 0.98 Identities = 13/40 (32%), Positives = 21/40 (52%), Gaps = 2/40 (5%) Query: 111 GNSDPRGFGHIGI--LVPDVDVACARFEQLGVKFIKRPND 148 GN D GFG++G+ + + + C +EQ+ V P D Sbjct: 126 GNDDDEGFGYLGLPPIAANEPILCLIWEQIAVLMTGSPED 165 >At1g15380.1 68414.m01841 lactoylglutathione lyase family protein / glyoxalase I family protein contains glyoxalase family protein domain, Pfam:PF00903 Length = 174 Score = 28.7 bits (61), Expect = 2.3 Identities = 14/50 (28%), Positives = 23/50 (46%), Gaps = 2/50 (4%) Query: 120 HIGILVPDVDVACARFEQLGVKFIKR--PNDGKMKGLAFIQDPDGYWIEI 167 HI + V + E++G+ +++ G F DPDG+ IEI Sbjct: 83 HISFQCESMGVVEKKLEEMGIDYVRALVEEGGIQVDQLFFHDPDGFMIEI 132 >At3g01810.1 68416.m00123 expressed protein Length = 921 Score = 28.3 bits (60), Expect = 3.0 Identities = 16/55 (29%), Positives = 27/55 (49%), Gaps = 3/55 (5%) Query: 69 YENPAEVPR--DDDARTKWAMT-RKATLELTYNWGTESDDSSYHNGNSDPRGFGH 120 Y AE+ DDD + ++T +TLE + +++ + N +PRG GH Sbjct: 194 YYKEAEIASITDDDISSHSSLTVSSSTLESNGGFSVRTEEEEHERINKNPRGNGH 248 >At3g27300.1 68416.m03412 glucose-6-phosphate 1-dehydrogenase / G6PD (ACG9) identical to glucose-6-phosphate 1-dehydrogenase (acg9) [Arabidopsis thaliana] GI:5732195 Length = 516 Score = 27.9 bits (59), Expect = 3.9 Identities = 19/41 (46%), Positives = 25/41 (60%), Gaps = 3/41 (7%) Query: 46 VLGMT--LLKQLHFPAMKFSLFFMGYENPAEVPRDDDARTK 84 VLG + L K+ FPA+ F+LF G+ NP EV AR+K Sbjct: 36 VLGASGDLAKKKTFPAL-FNLFHQGFLNPDEVHIFGYARSK 75 >At1g07645.1 68414.m00822 lactoylglutathione lyase family protein / glyoxalase I family protein Contains Pfam profile PF00903:glyoxalase family protein Length = 137 Score = 27.9 bits (59), Expect = 3.9 Identities = 13/43 (30%), Positives = 24/43 (55%), Gaps = 1/43 (2%) Query: 126 PDVDVACARFEQLGVKFIKRPNDGKM-KGLAFIQDPDGYWIEI 167 PDVD A R + G + + +P D + + + +++D DG + I Sbjct: 90 PDVDAAFKRAVENGAEAVSKPEDKEWGQKVGYVRDIDGIVVRI 132 >At5g40760.1 68418.m04946 glucose-6-phosphate 1-dehydrogenase / G6PD (ACG12) idential to glucose-6-phosphate 1-dehydrogenase (acg12) [Arabidopsis thaliana] GI:5732197 Length = 515 Score = 27.5 bits (58), Expect = 5.2 Identities = 19/41 (46%), Positives = 25/41 (60%), Gaps = 3/41 (7%) Query: 46 VLGMT--LLKQLHFPAMKFSLFFMGYENPAEVPRDDDARTK 84 VLG + L K+ FPA+ F+L+ G+ NP EV ARTK Sbjct: 36 VLGASGDLAKKKTFPAL-FNLYRQGFLNPDEVHIFGYARTK 75 >At5g03150.1 68418.m00263 zinc finger (C2H2 type) family protein contains Pfam domain, PF00096: Zinc finger, C2H2 type Length = 503 Score = 27.5 bits (58), Expect = 5.2 Identities = 11/32 (34%), Positives = 20/32 (62%) Query: 104 DDSSYHNGNSDPRGFGHIGILVPDVDVACARF 135 ++S+ N + P GF H G+ PD++ A ++F Sbjct: 236 NESNVMNNPNLPHGFVHRGVHHPDINAAISQF 267 >At4g21150.1 68417.m03057 ribophorin II (RPN2) family protein contains Pfam domain PF05817: Ribophorin II (RPN2) Length = 691 Score = 27.5 bits (58), Expect = 5.2 Identities = 16/51 (31%), Positives = 24/51 (47%), Gaps = 2/51 (3%) Query: 14 CQTPDPTTKDFMFQQTMYRIKDPRKTIPFYTGVLGMTLLKQLHFPAMKFSL 64 C+ + KD + Q +KD + + FY V G+ L K+ FP SL Sbjct: 105 CKIGEAGPKDIV-SQLQAGVKDAKLLLDFYYSVRGLVLAKE-QFPGTHISL 153 >At3g22330.1 68416.m02820 DEAD box RNA helicase, putative similar to RNA helicases GI:3775995, GI:3775987 from [Arabidopsis thaliana]; contains Pfam profiles PF00270: DEAD/DEAH box helicase, PF00271: Helicase conserved C-terminal domain Length = 616 Score = 27.5 bits (58), Expect = 5.2 Identities = 11/31 (35%), Positives = 17/31 (54%) Query: 97 YNWGTESDDSSYHNGNSDPRGFGHIGILVPD 127 Y G S+ ++G+SD GFG G+ P+ Sbjct: 525 YGGGRSGGSSNRYSGDSDRSGFGSFGMRSPE 555 >At3g06640.1 68416.m00772 protein kinase family protein contains Serine/Threonine protein kinases active-site signature, PROSITE:PS00108 Length = 763 Score = 26.6 bits (56), Expect = 9.1 Identities = 12/24 (50%), Positives = 13/24 (54%) Query: 147 NDGKMKGLAFIQDPDGYWIEIFTS 170 ND K FI DGY I +FTS Sbjct: 310 NDDKFSSEPFIDSSDGYPITLFTS 333 Database: arabidopsis Posted date: Oct 3, 2007 3:31 PM Number of letters in database: 12,070,560 Number of sequences in database: 28,952 Lambda K H 0.321 0.138 0.434 Gapped Lambda K H 0.279 0.0580 0.190 Matrix: BLOSUM62 Gap Penalties: Existence: 9, Extension: 2 Number of Hits to DB: 4,695,147 Number of Sequences: 28952 Number of extensions: 197077 Number of successful extensions: 403 Number of sequences better than 10.0: 19 Number of HSP's better than 10.0 without gapping: 9 Number of HSP's successfully gapped in prelim test: 10 Number of HSP's that attempted gapping in prelim test: 376 Number of HSP's gapped (non-prelim): 30 length of query: 173 length of database: 12,070,560 effective HSP length: 76 effective length of query: 97 effective length of database: 9,870,208 effective search space: 957410176 effective search space used: 957410176 T: 11 A: 40 X1: 16 ( 7.4 bits) X2: 37 (14.9 bits) X3: 62 (25.0 bits) S1: 41 (21.9 bits) S2: 56 (26.6 bits)
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